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Enlightenment is a Prophecy from Rising Sun. Once it is activated, it enables Treasure cards to be used as cantrips during your Action phase, allowing you to skip past your Treasure cards to try to put more useful cards into your hand. Be careful, though—this overrules any other effects that might otherwise enable you to play a Treasure in your Action phase for its normal ability!
Treasures can still be played in the Buy phase to do what they normally do, but if played in the Action phase, they produce +1 Card and +1 Action rather than everything they normally do.
You can turn these Treasures sideways to remind yourself that they didn't make .
Since Treasures are Actions, they can be used with Ways (from Menagerie) to get something other than +1 Card and +1 Action.
Highwayman (from Allies) can't stop your first Treasure from being used in an Action phase for +1 Card and +1 Action.
Other rules clarifications
Enlightenment even applies to Treasure cards that are already Actions; if you play Crown, Coronet, or an Action affected by Capitalism in the Action phase, you still get +1 Card and +1 Action instead of the card's usual effect.
Enlightenment only affects cards and not piles. This means that the Copper pile is not an Action pile for e.g. Training or Populate.
Treasures still remain as Actions when scoring (for e.g. Vineyard).
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Treasures are also Actions. When you play a Treasure in an Action phase, instead of following its instructions, +1 Card and +1 Action.